ejss

Eurasian Journal of Soil Science

2147-4249

Türkiye Toprak Bilimi Derneği

10.18393/ejss.1891362

Soil Sciences and Plant Nutrition (Other)

Toprak Bilimleri ve Bitki Besleme (Diğer)

Bacterial population and several soil properties across different vegetation stands in Andisols profiles at Mount Merbabu National Park, Indonesia

https://orcid.org/0009-0004-1720-0524

Suyana

Jaka

Department of Soil Science, Faculty of Agriculture, Sebelas Maret University, Indonesia

https://orcid.org/0009-0003-9589-1386

Tuada

Hayu Wisesa

Department of Soil Science, Faculty of Agriculture, Sebelas Maret University, Indonesia

https://orcid.org/0009-0000-3790-9814

Prasetyo

Dhany Eko

Department of Soil Science, Faculty of Agriculture, Sebelas Maret University, Indonesia

https://orcid.org/0000-0002-8473-9480

Muliawati

Endang Setia

Department of Agrotechnology, Faculty of Agriculture, Sebelas Maret University, Indonesia

https://orcid.org/0000-0001-7704-0754

Komariah

Department of Soil Science, Faculty of Agriculture, Sebelas Maret University, Indonesia

04 01 2026

15 2 253 266 10 07 2025 02 03 2026

2012

Eurasian Journal of Soil Science

A fundamental understanding of the forest stands and upland farming practices is imperative for effective land management, as it impacts the several soil properties. This research was to evaluate the relationship of bacterial population and several soil properties across different vegetation stands in Andisols soil profiles. Soil sampling was conducted on Mount Merbabu National Park (puspa (Schima noronhae Theaceae) and pine (Pinus merkusii Pinaceae)) and upland farming. For each vegetation observation plot unit, a pedon was created and 3 replications were carried out; and each pedon was sampled at soil profile depths 0-100 cm (0-10, 10-20, 20-30, 30-50, 50-70, and 70-100 cm) so 54 soil samples were obtained. The soil samples were then analyzed: bacterial population by the Spread Plate Count method, soil pH by pH meter, soil organic matter (SOM) by Walkley and Black method, and soil moisture by gravimetric method. The results showed that puspa had the highest means bacterial population (19.9 ± 2.67 ×10⁷ CFU g⁻¹), SOM (8.24 ± 1.06 %) and soil moisture (29.71 ± 1.93 %) which were significantly different from pine and upland farming; the highest bacterial population and SOM content in each stands was produced in the top layer (0-10 cm) and decreases with increasing soil depth; the lowest soil moisture was obtained in the top layer and increases with increasing soil depth; and bacterial population had a very significant relationship and are positively correlated with SOM and negatively correlated with soil moisture.

Bacterial population forest stands soil properties upland farming.

Sebelas Maret University for the Fundamental Research Grant

228/UN27.22/PT.01.03/2023

Abdila, A., Japarang, N., Agustin, N., Hafni, W., Annisi, A.D., Karim, H., Jumadi, O., 2022. Soil microorganism population in corn fields upon application of polyacrylate fertilizers. Indonesian Journal of Agricultural Sciences 27(1): 8-21 [in Indonesian].

Alfaro, F.D., Manzano, M., Marquet, P.A., Gaxiola, A., 2017. Microbial communities in soil chronosequences with distinct parent material: the effect of soil pH and litter quality. Journal of Ecology 105(6): 1709-1722.

Alfiyah, F., Nugroho, Y., Rudy, G.S. 2020. The Effect of Slope class and land cover on soil solum depth, effective root depth, and soil pH. Jurnal Sylva Scienteae 3(3): 499–508 [in Indonesian].

Alsabhan, A.H., Perveen, K., Alwadi, A.S., 2021. Heavy metal content and microbial population in the soil of Riyadh Region, Saudi Arabia. Journal of King Saud University - Science 34: 101671.

Aminudin, Y., Lestari, P., Prasetyo, E., Utomo, S., 2021. The abundance of soil macrofauna on the Mount Merapi post-eruption in Mount Merapi National Park area. Gorontalo Journal of Forestry Research 4(2): 98-112 [in Indonesian].

Anggraeni, R., Mahrup, M., Kusnarta, I. G. M., Silawibawa, P. 2022. Variation of soil moisture and temperature regime in land with low cloud cover based on Terra Modis Map in Lombok Island. Journal of Soil Quality and Management 1(1): 7-15 [in Indonesian].

Ayu, I.W., Oklima, A.M., Safitri, D., 2022. Analysis of soil moisture availability for corn and peanut planting times in dry fields in Berora Village, Lopok Subdistrict. Agrotechnology Journal 2(2): 1–14.

Bahram, M., Hildebrand, F., Forslund, S. K., Anderson, J. L., Soudzilovskaia, N. A., Bodegom, P. M., Bengtsson-Palme, J., Anslan, S., Coelho, L. P., Harend, H., Huerta-Cepas, J., Medema, M. H., Maltz, M. R., Mundra, S., Olsson, P. A., Pent, M., Põlme, S., Sunagawa, S., Ryberg, M., Tedersoo, L., Bork, P., 2018. Structure and function of the global topsoil microbiome. Nature 560(7717): 233–237.

Baso, M. S. G., Hasanah, U., Monde, A., 2014. Variability of soil physical properties and C organic in forest land and cocoa plantation (Theobroma cacao L.) in Sejahtera Village, Palolo Sub-district, Sigi Regency. E-Journal Agrotekbis 2(6): 565–572.

Bickel, S., Or, D., 2020. Soil bacterial diversity mediated by microscale aqueous-phase processes across biomes. Nature Communications 11(1): 116.

Bidkhani, N.O.G., Mobasheri, M.R., 2018. Influence of soil texture on the estimation of bare soil moisture content using MODIS images. European Journal of Remote Sensing 51(1): 911-920.

Brooks, K.N., Ffolliott, P.F., Magner, J.A., 2012. Hydrology and the Management of Watersheds. John Wiley & Sons. 533p.

Cheng, J., Zhao, M., Cong, J., Qi, Q., Xiao, Y., Cong, W., Deng, Y., Zhou, J., Zhang, Y., 2020. Soil pH exerts stronger impacts than vegetation type and plant diversity on soil bacterial community composition in subtropical broad-leaved forests. Plant and Soil 450(1): 273–286.

Choudhary, M., Datta, A., Jat, H.S., Yadav, A.K., Gathala, M.K., Sapkota, T.B., Das, A.K., Sharma, P.C., Jat, M.L., Singh, R., Ladha, J.K., 2018. Changes in soil biology under conservation agriculture based sustainable intensification of cereal systems in Indo-Gangetic Plains. Geoderma 313:193−204.

Delgado-Baquerizo, M., Oliverio, A.M., Brewer, T.E., Benavent-González, A., Eldridge, D.J., Bardgett, R.D., Maestre, F.T., Singh, B.K., Fierer, N., 2018. A global atlas of the dominant bacteria found in soil. Science 359(6373): 320–325.

Ebrahimi, M., Sinegani, A.A.S., Sarikhani, M.R., Mohammadi, S.A., 2017. Comparison of artificial neural network and multivariate regression models for prediction of Azotobacteria population in soil under different land uses. Computers and Electronics in Agriculture 140: 409-421.

El Mouridi, Z., Ziri, R., Douaik, A., Bennani, S., Lembaid, I., Bouharou, L., Brhadda, N., Moussadek, R., 2023. Comparison between Walkley-Black and Loss on Ignition methods for organic matter estimation in different Moroccan Soils. Ecological Engineering & Environmental Technology 24(4): 253-259.

Fadillah, N., Utomo, M., Afrianti, N. A., Sarno, S., 2022. Changes in soil chemical properties in soil profile due to the application of long-term N clearance system and fertilization on corn (Zea mays L.) plantations in Lampung State Polytechnic Experimental Garden. Journal of Agrotech Tropica 10(4): 627-632 [in Indonesian].

Fajar, I., Perwira, I.Y., Ernawati, N.M., 2022. The effect of acidity level (pH) on the growth of hexavalent chromium-tolerant bacteria from mangrove sediments in the Tukad Mati Estuary, Bali. Current Trends in Aquatic Science 5(1): 1–6 [in Indonesian].

Gou, Z., Zhai, Z., Zhang, Q., Li, Y., Feng, X., Zhang, Y., Zhao, X., Ding, M., 2024. Characteristics of organic carbon in tobacco-growing soils with different pH. Bangladesh Journal of Botany 53(3): 745-755.

Gunina, A., Kuzyakov, Y., 2021. Fromenergy to (soil organic) matter. Global Change Biology 28(1): 2169-2182.

Han, W.J., Wang, G.M., Liu, J.L., Ni, J., 2021. Effects of vegetation type, season, and soil properties on soil microbial community in subtropical forests. Applied Soil Ecology 158: 103813.

Hao, J., Chai, Y.N., Lopes, L.D., Ordóñez, R.A., Wright, E.E., Archontoulis. S., Schachtman, D.P., 2021. The effects of soil depth on the structure of microbial communities in agricultural soils in Iowa (United States). Applied and Environmental Microbiology 87: e02673-20.

Henny, H., Hasriati, N., Ridwan, M., 2023. Andisol soil properties and feasibility of potato farming with tractor tillage and organic fertilizer in Kerinci Regency. Journal of Tropical Agricultural Engineering and Biosystems 11(1): 63-72 [in Indonesian].

Hermans, S.M., Buckley, H.L., Case, B.S., Curran-Cournane, F., Taylor, M., Lear, G., 2017. Bacteria as emerging indicators of soil condition. Applied and Environmental Microbiology 83: e02826 16.

Hermans, S. M., Buckley, H. L., Case, B. S., Curran-Cournane, F., Taylor, M., Lear, G., 2020. Using soil bacterial communities to predict physico-chemical variables and soil quality. Microbiome 8(1): 79.

Hu, L., Wang, X., Song, X., Dai, D., Ding, L., Degen, A. A., Wang, C., 2023. Vertical patterns of soil bacterial and fungal communities along a soil depth gradient in a natural picea crassifolia forest in Qinghai Province, China. Forests 14(5): 1016.

Huang, C., He, Y., Zhou, L., Liu, R., Chen, H., Du, Z., Fu, Y., Zhu, Y., Zhou, Y., Wu, C., Zhou, X., 2024. Opposite effects of soil pH on bacteria and fungi β diversity in forests at a continental scale. Journal of Environmental Management 370: 122428.

Hutamy, H.B., Nuraini, Y., 2019. Diversity and population of phosphate solubilizing bacteria in various land uses at UB Forest. Jurnal Tanah dan Sumberdaya Lahan 6(1): 1113-1122 [in Indonesian].

Indrajaya, Y., Handayani, W., 2008. Potency of Merkus Pine (Pinus merkusii Jungh. et de Vriese) forest as landslide control in Java. Forest Info 5(3): 231-240.

Irfan, M., 2014. Isolation and enumeration of peat soil bacteria in PT Tambang Hijau's Oil Palm Plantation, Tambang District, Kampar Regency. Agrotechnology Journal 5(1): 1–8 [in Indonesian].

Islamzade, T., İslamzade, R., Azizov, R., Babayeva, T., Aliyeva, A., Haciyeva, X., Ashurova, N., 2025. Impact of Cadmium-contaminated water and irrigation levels on microbiological properties of soils with different textures. Eurasian Journal of Soil Science 14(2): 107-115.

Istiqomah, N., Indarto, T. N., Nugroho, V. A., Prayogo, C., 2017. Nitrogen availability and soil bacteria population under fertilization treatments in the System of Rice Intensification. Indonesian Soil and Climate Journal 41(2): 1-10.

Jiang, P., Wan, X., Che, M., Li, L., Liu, M., 2025. Soil depth exerts greater effect on bacterial community than spatial structure in Longmenshan fault zone. Applied and Environmental Microbiology 91: e01161-24.

Joshi, R.K., Garkoti, S.C., 2023. Influence of vegetation types on soil physical and chemical properties, microbial biomass and stoichiometry in the central Himalaya. Catena 222: 106835.

Kaiser, K., Wemheuer, B., Korolkow, V., Wemheuer, F., Nacke, H., Schöning, I., Schrumpf, M., Daniel, R., 2016. Driving forces of soil bacterial community structure, diversity, and function in temperate grasslands and forests. Scientific Reports 6(1): 33696.

Kaur, M., Li, J., Zhang, P., Yang, H-f., Wang, L., Xu, M., 2022. Agricultural soil physico- chemical parameters and microbial abundance and diversity under long-run farming practices: A greenhouse study. Frontiers in Ecology and Evolution 10: 1026771.

Kim, H.S., Lee, S.H., Jo, H.Y., Finneran, K.T., Kwon, M.J., 2021. Diversity and composition of soil Acidobacteria and Proteobacteria communities as a bacterial indicator of past land-use change from forest to farmland. Science of the Total Environment 797: 148944.

Li, W., Kuzyakov, Y., Zheng, Y., Li, P., Li, G., Liu, M., Alharbi H.A., Li, Z., 2022. Depth effects on bacterial community assembly processes in paddy soils. Soil Biology and Biochemistry 165: 108517.

Liu, J., Gul, Wazir Z., Hou, G-Q., Wang, G-Z., Rong, F-X., Xu, Y-Z., Liu, K., Li, M-Y., Liu, A-J., Liu H-L., 2023. The dependent correlation between soil multifunctionality and bacterial community across different farmland soils. Frontiers in Microbiology 14:1144823.

Lladó, S., López-Mondéjar, R., Baldrian, P., 2017. Forest soil bacteria: diversity, involvement in ecosystem processes, and response to global change. Microbiology and Molecular Biology Reviews 81: e00063-16.

Maturin, L., Peeler, J. T., 2001. BAM: Aerobic plate count. US Food and Drug Administration: Silver Spring, MD, USA.

Mayer, M., Prescott, C. E., Abaker, W. E. A., Augusto, L., Cécillon, L., Ferreira, G. W. D., James, J., Jandl, R., Katzensteiner, K., Laclau, J. P., Laganière, J., Nouvellon, Y., Paré, D., Stanturf, J. A., Vanguelova, E. I., Vesterdal, L., 2020. Influence of forest management activities on soil organic carbon stocks: A knowledge synthesis. Forest Ecology and Management 466: 118127.

Menyailo, O.V., Sobachkin, R.S., Makarov, M.I., Cheng, C.-H., 2022. Tree species and stand density: The effects on soil organic matter contents, decomposability and susceptibility to microbial priming. Forests 13: 284.

Muchane, M. N., Sileshi, G. W., Gripenberg, S., Jonsson, M., Pumariño, L., Barrios, E., 2020. Agroforestry boosts soil health in the humid and sub-humid tropics: A Meta-Analysis. Agriculture, Ecosystems and Environment 295: 106899.

Mundra, S., Kjonaas, O. J., Margado, L. N., Krabberod, A. K., Ransedokken, Y., Kauserud, H., 2021. Soil depthmatters: shift in composition andinter-kingdom co-occurrence patternsof microorganisms in forest soils. FEMS Microbiology Ecology 97(3): 1-14.

Nadal-Romero, E., Otal-Laín, I., Lasanta, T., Sánchez-Navarrete, P., Errea, P., Cammeraat, E., 2018. Woody encroachment and soil carbon stocks in subalpine areas in the Central Spanish Pyrenees. Science of the Total Environment 636: 727–736.

Nelson, D.W., Sommers, L.E., 1996. Total carbon organic carbon and organic matter. In: Methods of Soil Analysis: Part 3 Chemical Methods, 5.3. Sparks, D.L. Page, A.L., Helmke, P.A., Loeppert, R.H., Soltanpour, P.N., Tabatabai, M.A., Johnston, C.T., Sumner, M.E. (Eds.). SSSA Book Series No. 5. ASA-SSSA Madison WI, USA, pp. 961–1010.

Niu, Y., Li, X., Wang, C., Han, Y., Wang, Z., Yang, J., 2024. Contrasting altitudinal patterns and composition of soil bacterial communities along stand types in Larix principis-rupprechtii Forests in Northern China. Forests 15: 392.

Novara, A., Pisciotta, A., Minacapilli, M., Maltese, A., Capodici, F., Cerdà, A., Gristina, L., 2018. The impact of soil erosion on soil fertility and vine vigor. A multidisciplinary approach based on field, laboratory and remote sensing approaches. Science of the Total Environment 622–623: 474–480.

Novak, V., Hlavacikova, H., 2019. Soil-water content and its measurement. In: Applied Soil Hydrology. Theory and Applications of Transport in Porous Media. Hassanizadeh, S.M. (Ed.). vol 32. Springer, Cham. pp. 49-61.

Noviana, L., Raharjo, B., 2009. Viability of rhizobacteria Bacillus sp. DUCC-BR-K1.3 in peat soil substrate substituted with solid waste from the cigarette industry. Bioma : Berkala Ilmiah Biologi 11(1): 30-39 [in Indonesian].

Pepper, I.L., Gerba, C.P., 2004. Environmental microbiology: a laboratory manual. London: Elsevier academic press.

Portell, X., Pot, V., Garnier, P., Otten, W., Baveye, P.C., 2018. Microscale heterogeneity of the spatial distribution of organic matter can promote bacterial biodiversity in soils: insights from computer simulations. Frontiers in Microbiology 9:1583.

Prijono, S., Laksamana, M.T.S., 2016. Study of transpiration rates of Peltophorum dassyrachis and Gliricidia sepium in hedge plant cultivation systems and their effect on unsaturated hydraulic conductivity. J-PAL (Indonesian Journal of Environment and Suistainable Development) 7(1): 15–24 [in Indonesian].

Priyadi, P., Kurniawati, N., Nugroho, P. A., 2018. Biological activity of rubber plantation-derived soil at various moisture conditions. Journal of EnviScience (Environment Science) 2(1): 10-15 [in Indonesian].

Qi, J.Y., Wang, X., Zhao, X., Pu, C., Kan, Z.R., Li, C., Liu, P., Xiao, X.P., Lal, R., Zhang, H.L., 2019. Temporal variability of soil organic carbon in paddies during 13-year conservation tillage. Land Degradation and Development 30(15): 1840–1850.

Rayment, G.E., Lyons, D.J., 2011. Soil chemical methods: Australasia. CSIRO publishing. 495p.

Ritonga, A.G., Rauf, A., Jamilah., 2016. The characteristics of soil biology in various land use in Petani Watershed Deli Serdang Regency Sumatera Utara. Journal of Agroecotechnology Universitas Sumatera Utara 4(3): 108037 [in Indonesian].

Roose, E.J., Lal, R., Feller, C., Barthes, B., Stewart, B.A., 2006. Soil Erosion and Carbon Dynamics. CRC Press. 378p.

Rosalina, F., Maipauw, N. J., 2019. Soil chemical properties in some vegetation types. Journal Median 11(1): 1-9 [in Indonesian].

Staszel-Szlachta, K., Błońska, E., Lasota, J., 2025. Stabilization of soil organic matter in Luvisols under the influence of various tree species in temperate forests. Scientific Reports 15(1): 1286.

Saibi, N., Tolangara, A.R., 2017. Decomposition of Avecennia lanata litter at various soil depths. Techno: Jurnal Penelitian 6(01): 55-63 [in Indonesian].

Siahaan, F.A., Irawanto, R., Rahadiantoro, A., Abiwijaya, I.K., 2018. Topsoil properties under different vegetation in Purwodadi Botanical Garden. Indonesian Soil and Climate Journal 42(2): 91-98 [in Indonesian].

Soil Survey Staff, 2014. Keys to soil taxonomy. Government Printing Office, Washington. Available at [Access date: 07.10.2025]: https://nrcs.app.box.com/s/xi57bj6zyo601eokr7v715mkdpeaa81h/file/1147478400323

Solihin, M.A., Fitriatin, B.N., 2017. Soil microbe distribution in various types of land use in the North Bandung Area. Soilrens Journal 15(1): 38–45 [in Indonesian].

Song, L., Yang, T., Xia, S., Yin, Z., Liu, X., Li, S., Sun, R., Gao, H., Chu, H., Ma, C. 2022. Soil depth exerts stronger impact on bacterial community than elevation in subtropical forests of Huangshan Mountain. Science of the Total Environment 852: 158438.

Suyana, J., Komariah., Senge, M., 2010. Conservation techniques for soil erosion control in tobacco-based farming system at steep land areas of Progo Hulu Subwatershed, Central Java, Indonesia. International Journal of Agricultural and Biosystems Engineering 4(5): 287-294.

Suyana, J., Nugraheni., 2022. Effect of mulch and strengthened terrace strips on erosion, sediment enrichment ratio, and nutrient loss through erosion. Journal of Tropical Soils 27(3): 133-145.

Suyana, J., Krismonanto, W., Muliawati, E.S., Widijanto, H., Hartati, S., 2022a. The characteristics of soil organic carbon (SOC) at forest stands of Mount-Merbabu National Park and upland farming. IOP Conference Series: Earth and Environmental Science Volume 1114, 9th International Conference on Sustainable Agriculture and Environment 24/08/2022 - 25/08/2022. 1114: 012052.

Suyana, J., Krismonanto, W., Muliawati, E.S., Widijanto, H., 2022b. Characteristics of vegetation, nutrient nitrogen and soil organic carbon in forest stands of Mount Merbabu National Park and upland farming. Journal of Watershed Management Research 6(2): 141-160.

Suyana, J., Prasetyo, D.E., Fauziah, S., Muliawati, E.S., Cahyono, O. 2025. Fungi population and soil chemical and physical properties across different vegetation stands in andisol soil profiles. Eurasian Journal of Soil Science 14(4): 345-358.

Szypłowska, A., Lewandowski, A., Yagihara, S., Saito, H., Furuhata, K., Szerement, J., Kafarskia, M., Wilczeka, A., Majcherg, J., Woszczyka, A., Skierucha, W., 2021. Dielectric models for moisture determination of soils with variable organic matter content. Geoderma 401: 115288.

Thai, S., Pavlů, L., Vokurková, P., Thet, B., Vejvodová, K., Drábek, O., Tejnecký, V., 2024 Changes in the composition of soil organic matter after the transformation of natural beech stands into spruce monoculture. Soil Systems 8: 74.

Upton, R.N., Sielaff, A.C., Hofmockel, K.S., Xu, X., Polley, H., Wilsey, B.J., 2020. Soil depth and grassland origin cooperatively shape microbial community co‐occurrence and function. Ecosphere 11(1): 1-20.

van der Heijden, M.G.A., Bardgett, R.D., van Straalen, N.M., 2008. The unseen majority: soil microbes as drivers of plant diversity and productivity in terrestrial ecosystems. Ecology Letters 113: 296–310.

Visconti-Moreno, E.F., Valenzuela-Balcázar, I.G., 2023. Pores distribution influences the soil microorganism's response to changes in temperature and moisture. Eurasian Journal of Soil Science 12(1): 28-36.

Wan, W., Tan, J., Wang, Y., Qin, Y., He, H., Wu, H., Zuo, W., He, D. 2020. Responses of the rhizosphere bacterial community in acidic crop soil to pH: Changes in diversity, composition, interaction, and function. Science of the Total Environment 700: 134418.

Wang, Y., Chen, L., Xiang, W.H., Ouyang, S., Zhang, T.D., Zhang, X.L., Zeng, Y., Hu, Y., Luo, G., 2021. Forest conversion to plantations: a meta-analysis of consequences for soil and microbial properties and functions. Global Change Biology 27: 5643–5656.

Wang, C., Yu, Q.Y., Ji, N.N., Zheng, Y., Taylor, J.W., Guo, L.D., Gao, C., 2023. Bacterial genome size and gene functional diversity negatively correlate with taxonomic diversity along a pH gradient. Nature Communications 14(1): 7437.

Wu, H., Cui, H., Fu, C., Li, R., Qi, F., Liu, Z., Yang, G., Xiao, K., Qiao, M. 2024. Unveiling the crucial role of soil microorganisms in carbon cycling: A review. Science of the Total Environment 909: 168627.

Xiao, L., Zhou, S., Zhao, R., Greenwood, P., Kuhn, N. J., 2020. Evaluating soil organic carbon stock changes induced by no-tillage based on fixed depth and equivalent soil mass approaches. Agriculture, Ecosystems and Environment 300: 106982.

Yin, J., Chen, H., Duan, P., Zhu, K., Ma, Y., Xu, Y., Guo, J., Li, R., Chen, Q., 2023. Soil microbial communities as potential regulators of N2O sources in highly acidic soils. Soil Ecology Letters 5: 230178.

Zhang, Y. Y., Wu, W., Liu, H., 2019. Factors affecting variations of soil pH in different horizons in hilly regions. PLoS ONE 14(6): e0218563.

Zhang, J., Li, Y., Yang, T., Liu, D., Liu, X., Jiang, N. 2021. Spatiotemporal variation of moisture in rooted-soil. Catena 200: 105144.

Zhang, D., Zhao, Y., Qi, H., Shan, L., Chen, G., Ning, T., 2024. Effects of micro-topography and vegetation on soil moisture on fixed sand dunes in Tengger Desert, China. Plants 13(11): 1571.

Zhou, W., Han, G., Liu, M., Zeng, J., Liang, B., Liu, J., Qu, R., 2020. Determining the distribution and interaction of soil organic carbon, nitrogen, pH and texture in soil profiles: a case study in the Lancangjiang River Basin, Southwest China. Forests 11(5): 532.