No less than 66 species have been assigned to the genus Paralorryia, at least momentarily (subjective synonyms set apart). Based on its type species, Lorryia cumbrensis, and on the study of its phanerotaxy, the genus Paralorryia was synonymized with Calotydeus by André (2025). Besides cumbrensis, only 6 other species are assigned to Calotydeus. All other species are scattered across five genera: Brachytydeus (51 species), Neohomeotydeus (2 species), Nudilorryia (1 species), Ueckermannia (1 species) and Pretydeus (Pretydeinae) (1 species). 3 species are inquirendae and 1 is a nomen nudum. Nomenclatural changes are: Calotydeus andreae (new combination), Calotydeus costaricensis (n. comb.), Calotydeus nikitensis (n. comb.), Calotydeus spineus (n. comb.), Brachytydeus aegyptocitri (n. comb.) and Brachytydeus fodderi (n. comb.).
Not applicable.
This study was not supported by any foundation.
Many thanks to F.M. Momen (Egypt) for providing infor-mation on El-Bagoury and Paralorrya gizai. Many thanks also to E.A. Ueckermann (South Africa) for reviewing a preliminary draft of this paper and to the anonymous referees for their careful review of the manuscript.
| Primary Language | English |
|---|---|
| Subjects | Acarology |
| Journal Section | Research Article |
| Authors | |
| Submission Date | August 31, 2025 |
| Acceptance Date | October 1, 2025 |
| Publication Date | January 6, 2026 |
| Published in Issue | Year 2026 Issue: 2026 |
The aim of the journal Acarological Studies (AS) is to promote acarological research worldwide. The journal is international, peer reviewed, open access, on-line and it has no publication fees.
AS publishes high quality research papers on any aspect of mites, including anatomy, behaviour, biodiversity, conservation, biological control, ecology, medical acarology, morphology, phylogeny, physiology, systematics, veterinary acarology and zoogeography. The journal also invites the submission of case studies, monographs, revisions, reviews, catalogues, checklists, descriptions of taxa and nomenclature for consideration.
All manuscripts must be accompanied by a separate title and statement page (click here to download template file of the title and statement page).
This page should contain the full title. The title should be short and informative, and not contain non-standard acronyms or abbreviations or taxon's authors. The higher taxa containing the taxa dealt with in the manuscript should be indicated in parentheses in the title. The full names of all authors must be given and surname should be typed in the upper case. The address of each author should be given in italic each starting a separate line. Authors are strongly encouraged to include an ORCID with their manuscript submission or revision (for more information on how to register for an ORCID click here). The contact e-mail address should be provided for the clearly identified corresponding author.
Authors’ contributions to the manuscript, ethics approval, funding source(s), conflict of interest statements, and acknowledgements should be given here.
The manuscripts should be written in English or Turkish. There are no word limits for the manuscript in this journal. The manuscripts should be MS-Word processed, typed with size 12 in Times New Roman font, double-spaced, continuous line numbered, with 2.5 cm margins on each side. All pages should be numbered consecutively in the bottom. The manuscript should be compiled in the following order: title, abstract, keywords; introduction, material and methods, results and discussion, acknowledgments, references, tables with captions on individual pages and figure captions. The figures should be saved separately from the text. The file of main manuscript does not contain identification information about the authors such as name, e-mail address, affiliation and etc. (click here to dowload template file of the main text).
Abstract
The abstract should present the general scope, and give summarize information on the main results and conclusions of the paper. In the manuscripts written in Turkish, the abstract in English must be included in addition to the Turkish abstract. Any new names or new combinations proposed in the manuscript should be referred in the abstract. The abstract should preferably be kept below 350 words, and citations should generally be avoided where possible.
Keywords
Keywords are used for indexing purposes. For this reason, it is better to give keywords that are not present at the title. Abbreviations and acronyms should be avoided.
Introduction
The introduction should provide concisely description of the basic background, rationale, aims and objectives of the study. It should not include the findings.
General Information
The manuscripts should be written in English or Turkish. There are no word limits for the manuscript in this journal. The manuscripts should be MS-Word processed, typed with size 12 in Times New Roman font, double-spaced, continuous line numbered, with 2.5 cm margins on each side. All pages should be numbered consecutively in the bottom. The manuscript should be compiled in the following order: title, abstract, keywords; introduction, material and methods, results and discussion, acknowledgments, references, tables with captions on individual pages and figure captions. The figures should be saved separately from the text. The file of main manuscript does not contain identification information about the authors such as name, e-mail address, affiliation and etc.
Tables should be editable tables in a word document or main text. All columns must have headings. If a table continues on more than one page, repeat column headings on subsequent page(s). Use approved abbreviations already defined in the text and define them in the general footnote.
The figures should be saved separately from the text. They should be in their native format for best quality. The figures should be at least 600 dpi, or 1200 dpi for line graphs. The following file types of figures are accepted: png (preferred), tiff, gif or jpeg.
All tables and figures should have a caption or legend, they should be labelled, and the labels should be numbered unless there is only one table or figure. All tables and figures should be referred to in the text (e.g., “Table 1”, “Fig. 2”, “Figs 3-5”).
Citations should be arranged chronological and alphabetical within year. All references cited in the text must appear in the literature cited section, and all items in this section must be cited in the text. Citation of unpublished studies or reports is not permitted, i.e., a volume and page number must be available for serials and a publisher, city, state, and full pagination for books. Work may be cited as “in press” only when the paper has been accepted for publication. Personal communications do not appear in the literature-cited section.
Electronic Publication & Nomenclatural Information
Article 8.5.3. To be considered published, a work issued and distributed electronically must be registered in the Official Register of Zoological Nomenclature (ZooBank) and contain evidence in the work itself that such registration has occurred.
The requirements for electronic publications are that the work be registered in ZooBank before it is published. This is a recent change to the International Code of Zoological Nomenclature. The papers published in Zootaxa and Zookeys by International Commission on Zoological Nomenclature (2012) explain how to do this, to make the new names available. Here is the web site for registration of your paper in ZooBank: http://zoobank.org/. We remind you that the data entered in ZooBank are your responsibility and we thank you making sure that the information provided is accurate and complete.
Materials and Methods
In this section, it should be provided concise but complete information about the materials and the analytical and statistical procedures used. This part should be as clear as possible to enable other scientists to repeat the research presented. Brand names should be supplied for all mentioned equipment, instruments, consumables etc.
Authors are responsible for verifying whether collection permit is necessary in the country from which their specimens originate, and for timely acquisition of such a permit. DNA sequences must be deposited in a public database (e.g., Genbank) and accession numbers should be provided in the manuscript.
Results
In this section it should be presented only the data which support the objective without any discussion of the results. Figures and tables are used to present the results, but information in figures and tables should not be duplicated in the text.
Discussion
In this section the results are discussed and interpreted in the context of the existing knowledge. It can be presented here the relationships and generalizations shown by the results. Information in the Introduction and Results sections should not be repeated here. Main conclusion of the study can be clearly given in the end of this section.
Authors’ contributions
All authors should make substantial, meaningful contributions to the manuscript. In the title page and main manuscript text, the authors must include a statement outlining their individual contributions to the manuscript by using the relevant CRediT roles (Brand et al., 2015; https://doi.org/10.1087/20150211). The necessary CRediT roles are: Conceptualization; Data curation; Formal analysis; Funding acquisition; Investigation; Methodology; Project administration; Resources; Software; Supervision; Validation; Visualization; Writing - Original draft; Writing - Review & Editing.
Authors must meet at least three of the criteria, please click here for details of the criteria. When there are multiple people serving in the same role, a degree of contribution may optionally be specified as ‘lead’, ‘equal’, or ‘supporting’.
An example of an authors’ contribution statement using CRediT roles with degree of contribution:
Author 1: Conceptualization, data curation, formal analysis (lead), visualization (supporting), writing - original draft (lead). Author 2: Investigation, formal analysis (supporting), methodology (equal), visualization (lead), software, writing - original draft (supporting). Author 3: Project administration, supervision, methodology (equal), writing - review & editing.
It is recommended that corresponding authors assume responsibility for role assignment, and that all contributors be given the opportunity to review and confirm assigned roles.
A contribution statement is not required for a manuscript with a single author.
All contributors who do not meet the criteria for authorship should be listed in the section, ‘Acknowledgements’.
Statement of ethics approval
If the work involves the use of human subjects, the author(s) should ensure that the work described has been carried out in accordance with The Code of Ethics of the World Medical Association (Declaration of Helsinki, 2013). The author(s) should include a statement in the manuscript that informed consent was obtained for experimentation with human subjects. The privacy rights of human subjects must always be observed. In addition, the manuscript must include the name of the ethics committee that approved the study and the committee’s reference number/ID.
For work involving data of minors/children/infants, author(s) should confirm that statements of written informed consent from legally authorized representatives/parents/guardians are available; if verbal informed consent was obtained, reasons for this must be mentioned.
If the work involves animal experiments, it should be carried out in accordance with the relevant national institute of health guidelines for the care and use of laboratory animals and the author(s) should clearly state in the manuscript that such guidelines have been followed and also provide an approval number for the research protocol granted by an Animal Ethics Committee in accordance with international principles in this section.
If the manuscript does not report on or involve any human data or animal experiments, please state “Not applicable.” in this section.
If authors use AI tools for content generation, editing, coding and analytical assistance, this must be clearly stated in an appropriate section of the manuscript or in this section.
Funding
List here all funding details required by your funding and grant-awarding bodies. Names of funding organisations should be written in full.
Conflict of interest
The authors should declare whether they have conflict of interest or not.
Acknowledgements
It should be given in a separate section at the end of the manuscript before the references, and included them on the title page. It should be listed here those individuals who provided help during the research.
References
Citations should be arranged chronological and alphabetical within year. All references cited in the text must appear in the literature cited section, and all items in this section must be cited in the text. References should be arranged alphabetically. Citation of unpublished studies or reports is not permitted, i.e., a volume and page number must be available for serials and a publisher, city, state, and full pagination for books. Abstracts not subjected to peer review may not be cited in the text or in the literature cited section. Work may be cited as “in press” only when the paper has been accepted for publication. Personal communications do not appear in the literature-cited section.
Citation style in the text
One author
Fain (1998) or (Kontschán, 2020).
Two authors
Seeman and Baker (2013) or (Fan and Zhang, 2005).
Three and more authors
Lindquist et al. (2009) or (Khaustov et al., 2017).
(Grandjean, 1944; Düzgüneş, 1963; Khanjani, 2005; Keskin, 2016) Chronological.
(Ermilov, 2010, 2015; Bochkov and Oconnor, 2015) Chronological and alphabetical within the same year.
(Ueckermann et al., 2015a,b) As long as the first author and the year are the same, it should be used the letter designators to distinguish between publications.
Style of the literature cited section
Journal article, 1 author
Ewing, H.E. 1911. New predaceous and parasitic Acarina. Psyche, 18: 37-43.
Journal article, 2 authors
Fan, Q.-H. and Ueckermann, E.A. 2016. Resurrection of the genus Nonocaligus Habeeb with redefination of Nonocaligus and Mullederia Wood (Acari: Stigmaeidae). Systematic and Applied Acarology, 21 (11): 1447-1449.
Journal article, 3 and more authors
Cakmak, I., Janssen, A., Sabelis, M.W. and Baspinar, H. 2009. Biological control of an acarine pest by single and multiple natural enemies. Biological Control, 50: 60-65.
doi: 10.1016/j.biocontrol.2009.02.006
Note that titles of serial publications must be written in full. DOI (if available) should be given. If there are more than 10 authors, then include names of the first 10, followed by et al.
Electronic resource
OConnor, B. and Klimov, P. 2012. Family Cheyletidae Leach, 1815. Available from http://insects.ummz.lsa.umich.edu/beemites/Species_Accounts/Cheyletidae.htm (Last accessed: 19 May 2018).
Book
Krantz, G.W. and Walter, D.E. 2009. A manual of acarology. Third edition. Texas Tech University Press, Lubbock, Texas, USA, 807 pp.
Book Chapter
Kethley, J. 1990. Acarina: Prostigmata (Actinedida). In: Soil biology guide. Dindal, D.L. (Ed.). John Wiley and Sons, New York, USA, 667-756.
Thesis
Le Goff, G. 2011. Benefits of aggregation in Tetranychus urticae. Unpublished PhD Dissertation. Earth and Life Institute, Biodiversity Research Centre, Faculty of Sciences, Catholic University of Louvain, Louvain-la-Neuve, Belgium, 298 pp.
Congress
Sullivan, G.T and Ozman-Sullivan, S.K. 2018. Is a cataclysmic mass extinction of mite species occurring in the shadows of high profile insect and vertebrate extinctions? XV International Congress of Acarology, 2-8 September 2018, Antalya, Türkiye, p. 117.
Ethical Principles
Acarological Studies (AS) is committed to upholding the highest standards of publication ethics and expects all parties (authors, referees, editors and publisher) in the publishing process to bear the following ethical responsibilities.
In keeping with its commitment to best practices in academic publishing, AS strongly supports and adheres to the Principles of Transparency and Best Practice in Scholarly Publication adopted by the Committee on Publication Ethics (COPE), Directory of Open Access Journals (DOAJ), Open Access Scholarly Publishers Association (OASPA) and World Association of Medical Editors (WAME). Acarological Studies utilises the process flow sheets developed by COPE in collaboration with Springer Nature when dealing with cases of fraudulent manipulation of the publication process.
Authors’ Responsibilities
The persons who participate in the design of studies, data collection or the analysis or interpretation of the data in a study, or meaningfully contribute to arrangement of the content, are described as authors. Only people who meet these criteria can be included in the list of authors of a paper.
Before you undertake any research you should familiarize yourself with the full meaning and application of ethics in research. There are numerous resources available, including the following link: https://www.niehs.nih.gov/research/resources/bioethics/whatis/index.cfm
In addition, authors should:
AS utilises the process flow sheet prepared by COPE in situations potentially related to ethical problems. Please click here to access the flow sheet prepared by the committee.
Reviewers’ Responsibilities
After initial evaluation, the manuscript is reviewed by at least two independent, expert referees. If there are substantial inconsistencies between the referees, more reviewers are asked to review the manuscript. In every stage of the evaluation process, the double-blind peer review process, in which the names of the referees and the authors are not disclosed, is used.
Reviewers should:
Reviewers must follow COPE Ethical Guidelines for Peer Reviewers prepared by the COPE. Please click here to access the guidelines.
Editors’ Responsibilities
Editors should facilitate the sharing of scientific output with the target readership group by meeting the needs of both readers and the authors. In addition, they should record the opinions and ideas of authors, readers, reviewers and members of the editorial board to help improve journal processes. Editors should also adhere to the highest standards and best practices in academic publishing, and reconsider the journal’s processes in the light of new information and circumstances.
At AS, the chief editor has overall responsibility and the section editors appointed by the chief editor manage the peer review processes of the relevant manuscripts on behalf of the chief editor. The section editors are responsible for deciding whether manuscripts are published in the journal. The section editors should also provide feedback on the journal’s performance and have input to the selection of editorial board members.
Editors should:
All editors are obliged to follow the guidelines in the Code of Conduct and Best Practice Guidelines for Journal Editors developed by COPE. In this combined version of the documents, the mandatory Code of Conduct for Journal Editors standards are shown in regular script and with numbered clauses, and the more aspirational ‘Best Practice’ recommendations are shown in italics. In addition, you can find a handbook, which is the main reference guide for editors, translated to Turkish (click here) with the support of Trakya University and TÜBİTAK – ULAKBİM, and distributed to the editors of scientific journals in Türkiye.
Publisher’s Responsibilities
The publisher:
Readers’ Responsibilities
AS expects readers to bear the following responsibilities, even if they are not in the publishing process.
Readers:
Artificial Intelligence (AI) Use Policy
The primary purpose of the policy on the use artificial intelligence (AI) tools in academic processes is to preserve scientific accuracy, transparency and research ethics. This policy requires all stakeholders to use AI technologies exclusively in accordance with these principles and to disclose their use of such tools clearly when necessary.
Acarological Studies (AS) takes an ethical and responsible approach to the use of AI, placing human-centred values at the forefront. AS complies with the international standards of organisations such as The Council of Higher Education (YÖK), the Committee on Publication Ethics (COPE), World Association of Medical Editors (WAME), the United Nations Educational, Scientific and Cultural Organization (UNESCO), Wiley and Springer Nature regarding the use of AI in research and publishing processes. AS stays up to date with the rapid developments in AI technologies and reserves the right to update its policies as necessary.
The following sections outline the responsibilities and principles relating to the use of AI for different roles.
For Authors
Use of AI-Generated Data or Visuals
AI-based translation tools may be used only for support. It is incumbent upon the authors to ensure the accuracy of the produced text and to verify its final version. Tables and analytical outputs generated by AI may be used provided they are not presented directly as real data. The publication of videos and images, including figures and micrographs, generated by AI is not permitted, except for those specifically used in a paper related to AI. The use of fabricated or synthetic data generated by AI or other means in the production of scientific results is strictly prohibited.
Authors’ Responsibility for Scientific Content and Accuracy
Content produced by AI tools may contain factual inaccuracies, fabricated references or erroneous data. Therefore, it is fully the authors’ responsibility to verify all AI-generated outputs, correct them when necessary and confirm their scientific appropriateness. AI-assisted content generation cannot replace scientific evaluation or scientific judgement.
Confidentiality and Personal Data Security
Authors should not upload confidential data, personal information and private data from human subjects or unpublished research results to general-purpose online AI models. When such content must be processed, only institution-approved, secured and closed AI systems can be used.
Plagiarism and Originality Responsibility
Any texts created by AI must be checked for plagiarism. Authors are responsible for ensuring the originality, proper attribution and ethical use of AI-assisted content. AI outputs may not be directly copied into a manuscript. The author must review them and make any necessary changes to ensure they fit with the scientific framework.
AI Systems Not Listed in Authors
AI tools cannot be listed as authors on a manuscript. All scientific content, evaluations and conclusions in the manuscript are the responsibility of the human authors. Because AI-generated content does not meet international authorship criteria, AI tools can’t be authors.
Disclosure of AI Use
If authors use AI tools (e.g., ChatGPT, DeepSeek, Gemini) for content generation, editing, coding and analytical assistance, this must be clearly stated in an appropriate section of the manuscript (e.g., Methods) or in the section of ethics statement. The statement must include the tool name, version, purpose of use and usage rate. Disclosure is not required when AI tools are used solely for translation, simple language editing or cross-reference checking.
For Reviewers
Protection of Confidential Materials
Reviewers should not upload the manuscript under review —or any supplementary files, figures, tables, datasets or other confidential materials— to any AI system. This restriction applies to online large language models, cloud-based analytical tools and any platform that transmits content to external servers. In accordance with author confidentiality and double-blind review principles, no information obtained during peer review can be transferred to AI models.
Limited Use of AI When Preparing Review Reports
Reviewers can use AI tools solely to improve the linguistic clarity of the review report text that they themselves have written. AI tools cannot be used for scientific assessment, data interpretation or decision-making. Academic evaluations must rely entirely on the reviewer’s expertise.
Responsibility for Review Report Content
Suggestions generated by AI tools should not be accepted automatically; they must be checked and verified by the reviewer. Reviewers are responsible for the content, scientific accuracy and appropriateness of their reports. AI systems can’t replace the reviewer’s expert judgement.
Disclosure of AI Use in Review Reports
If a reviewer uses AI tools when preparing a review report, this should be disclosed to the editor when necessary.
Journal’s Right to Audit
If improper AI use is suspected in a review report, AS reserves the right to investigate, invalidate the report, appoint a new reviewer or request further clarification. These measures aim to ensure transparency and uphold ethical standards in the editorial process.
For Editors
Permissible Uses of AI Tools
Editors may use AI tools for mechanical tasks such as checking the suitability of manuscripts, reviewing formatting, conducting preliminary linguistic screening and drafting cover letters. However, scientific decisions such as desk rejection, returning manuscripts to authors, requesting revisions or acceptance/rejection decisions must rely solely on the editor’s academic judgement.
Insights produced through AI-assisted evaluations (e.g., potential plagiarism and linguistic issues) are only preliminary indicators. They cannot be used directly in decision-making without editor verification. AI tools are assistive in nature and cannot replace editorial expertise.
Data Storage and Confidentiality
When using AI, editors should adhere to the principles of transparency, accountability, and human oversight. Confidential editorial files, unpublished data and reviewer reports should not be uploaded to AI systems. Editors must ensure that any AI platform they use does not use uploaded information for training model or for third-party purposes. All systems must operate solely in accordance with confidentiality agreements and in non-training (closed) modes.
For Readers
Critical Evaluation
Readers should be aware that although AI-based analyses in scientific manuscripts have passed through editorial and peer review filters, it is still important to evaluate the results critically. AS may request disclosure statements from authors regarding AI contributions when necessary. Readers should be aware that AI may be used for formatting, statistical summarization or content refinement, but scientific data, interpretations and conclusions are solely the authors’ responsibility.
Respect for Confidentiality and Copyright
Readers should avoid uploading published papers or datasets in bulk to AI tools for reprocessing, as such actions may violate copyright and confidentiality. Before using published content for model training or commercial data generation, users must consider applicable licensing terms.
For Data Repositories and Libraries
AI-Assisted Metadata and Archival Processing
Data repositories and libraries can use AI tools for tasks such as metadata verification, content classification and archival processing. However, confidential or unpublished materials, as well as user-specific private data cannot be used for AI model training.
Institutional archives and libraries should offer licensing options to prevent papers, datasets or other uploaded materials from being used automatically in AI model training. All AI-based operations and outputs must be subject to human oversight and must not be finalized without human approval.
AI in the Context of Licensing and Copyright
Libraries must clearly and transparently communicate in their policy documents the legal boundaries of AI tools regarding access to licensed resources, text and data mining, bulk data extraction and commercial uses.
Responsible Use of AI-Based Search Engines
When providing AI-based search services, libraries must offer transparent information regarding the model’s training sources, data privacy safeguards, content filtering (e.g., risks of generating inaccurate or fabricated information —hallucinations) and reliability scoring.
Publication Policy
Acarological Studies (AS) is a peer-reviewed, international scientific journal that publishes and promotes research in acarology worldwide. It is a fully open access journal, published continuously by DergiPark for the “Turkish Academic Network and Information Center (ULAKBİM)”, a unit of “The Scientific and Technological Research Council of Türkiye (TÜBİTAK)”. DergiPark provides online hosting services and an editorial workflow management system for academic journals published in Türkiye. The DergiPark project, undertaken by TÜBİTAK – ULAKBİM, is a public service in Türkiye whose mission is the web-based publication of academic journals and the operation of an online journal management system.
All manuscripts submitted for publication in AS will be pre-reviewed by the editor. Manuscripts that fall outside the aims and scope of the journal, are inadequate in terms of language and writing, not original or not sufficient for the requirements of a scientific study, will be rejected or may have revisions requested at this initial stage.
After initial evaluation, the manuscript will be double blind, peer reviewed by at least two independent, expert referees. If there are inconsistencies between the referees, more reviewers will be asked to review the manuscript. The editor will decide on the acceptance or rejection of the manuscript according to the reports of these referees. However, a manuscript may be rejected without external review by the editor if it is considered unacceptable, for example, a manuscript will be rejected without peer review if it does not comply with the instructions for authors or if it is beyond the scope of the journal.
AS uses a plagiarism checker to detect potential plagiarism in the manuscript. If the editor, editorial board member or reviewer detects plagiarism at any stage of the evaluation process, the corresponding author will be immediately notified that the manuscript has been refused. AS utilises the process flow sheets developed by COPE when dealing with cases of potential plagiarism in a submitted manuscript or a published article.
There is almost always similarity between published papers and a new manuscript that cites them but the similarity has to be limited and the works have to be correctly referenced.
The names and e-mail addresses entered in the journal system DergiPark will be used exclusively for the stated purposes of this journal and will not be made available for any other purpose or to any other party. Responsibility for technical content and for protection of proprietary material rests solely with the author(s) and their organizations and is not the responsibility of the publisher, the journal or its editorial staff.
AS provides full open access to its publications according to the principle that providing scientific research to the public free of charge increases global information sharing. There is no publication fee in AS. The journal does not charge authors any fee for submitting, processing or open access publishing. All published papers in AS are freely available online.
Acarological Studies provides full open access to its publications according to the principle that providing scientific research to the public free of charge increases global information sharing. There is no publication fee in Acarological Studies. The journal does not charge authors any fee for submitting, processing or open access publishing. All papers published in Acarological Studies are freely available online.
Acarological Studies is licensed under a Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International.
International Scientific Research Journal on Acarology