Dismantling of the genus Paralorryia (Acariformes: Tydeoidea: Tydeidae)

Abstract

No less than 66 species have been assigned to the genus Paralorryia, at least momentarily (subjective synonyms set apart). Based on its type species, Lorryia cumbrensis, and on the study of its phanerotaxy, the genus Paralorryia was synonymized with Calotydeus by André (2025). Besides cumbrensis, only 6 other species are assigned to Calotydeus. All other species are scattered across five genera: Brachytydeus (51 species), Neohomeotydeus (2 species), Nudilorryia (1 species), Ueckermannia (1 species) and Pretydeus (Pretydeinae) (1 species). 3 species are inquirendae and 1 is a nomen nudum. Nomenclatural changes are: Calotydeus andreae (new combination), Calotydeus costaricensis (n. comb.), Calotydeus nikitensis (n. comb.), Calotydeus spineus (n. comb.), Brachytydeus aegyptocitri (n. comb.) and Brachytydeus fodderi (n. comb.).

Keywords

• Tydeinae
• Brachytydeus
• Calotydeus
• nomenclature
• taxonomy

References

1. Akimov, I.A., Kolodochka, L.A., Pavlitshenko, P.G., Voitenko, A.N., Kulczycki, A.G., Vinnik, E. N. and Pogrebnyak, S.G. 1993. Industrial orchard mite associations in Ukraine and their structural peculiarities. Vestnik Zoologii, 6: 48-56. [In Russian]
2. Alabdulla, J. and Kanouh, M. 2023. First record of the mite Brachytydeus mali Oudemans (Acari: Tydeidae) on apple trees in southern Syria. Arab and Near East Plant Protection Newsletter, 89: 8. [Part in Arabic]
3. Ali, E., Kandeel, M.M.H., Abd-ElKaream, S. and El-Bassiony, M.N. 2016. Two new acarofauna inhabiting some fruit trees in north Sinai, Egypt (Acari: Cheyletidae & Tydeidae). Sinai Journal of Applied Sciences, 5 (3): 393-398. doi: 10.21608/sinjas.2016.78661
4. Alizade, G.A. 2019. The predatory actinedid mites (Acariformes: Actinedida) of Khachmaz region of Azerbaijan. Journal of Entomology and Zoology Studies, 7 (1): 155-159.
5. André, H.M. 1980. A generic revision of the family Tydeidae (Acari: Actinedida). IV. Generic descriptions, keys and conclusions. Bulletin et Annales de la Socie ́te ́ royale belge d’Entomologie, 116: 103-130, 139-168.
6. André, H.M. 1984. Tydeinae (Acari: Tydeidae) from Belgium. I. The genus Homeotydeus. Bulletin et Annales de la Société Royale Belge d’Entomologie, 120: 117-122.
7. André, H.M. 2005. In search of the true Tydeus (Acari: Tydeidae). Journal of Natural History, 39: 975-1001. doi: 10.1080/00222930400002838
8. André, H.M. 2025. Sixty years after "A review of the genera of the family Tydeidae" (Acariformes: Tydeoidea: Tydeidae). Acarologia, 65 (1): 149-172. doi: 10.24349/l5vb-767l

9. Araújo, W.S. de and Daud, R.D. 2017. Insights on plant mite occurrence in natural vegetation remnants from Brazil. Systematic and Applied Acarology, 22: 302-323.
10. Baker, E.W. 1943. Nuevos Tydeidae Mexicanos (Acarina). Revista de la Sociedad Mexicana de Historia Natural, 4: 181-190. [In Spanish]
11. Baker, E.W. 1944. Seis especies de Lorryia (Acarina, Tydeidae). Anales del Instituto de Biológia de la Universidad Nacional de México, 15 (1): 215-222. [In Spanish]
12. Baker, E.W. 1947. Notes on mites of the family Tydeidae (Acarina) with descriptions of two new species. Proceedings of the Entomological Society of Washington, 49: 133-136.
13. Baker, E.W. 1965. A review of the genera of the family Tydeidae (Acarina). In: Advances in acarology 2. Naegele, J.A. (Ed.) Cornell University Press, Ithaca, NY, USA, 96-133.
14. Baker, E.W. 1968a. The genus Lorryia. Annals of the Entomological Society of America, 61: 986-1008. doi: 10.1093/aesa/61.4.986
15. Baker, E.W. 1968b. The genus Paralorryia. Annals of the Entomological Society of America, 61: 1097-1106. doi: 10.1093/aesa/61.5.1097
16. Baker, E.W. 1970. The genus Tydeus: subgenera and species group with descriptions of new species (Acarina: Tydeidae). Annals of the Entomological Society of America, 63: 163-177. doi: 10.1093/aesa/63.1.163
17. Bernini, F., Castagnoli, M. and Nannelli, R. 1995. Arachnida Acari. In: Checklist delle specie della fauna italiana. Minelli, A., Ruffo, S. and La Posta, S. (Eds). Commission of the European Communities, 24: 24-66. [In Italian]
18. Bozai, J. 1997. Tydeidae fauna data from Hungary (Acari, Tydeidae). Növényvédelem, 33: 77-79. [In Hungarian]
19. Castagnoli, M. 1984. Contributo alla conoscenza dei Tideidi (Acarina: Tydeidae) delle piante coltivate in Italia. Redia, 67: 307-322. [In Italian]
20. Cooreman, J. 1958. Notes et observations sur les acariens. VII. Photia graeca n. sp. (Acaridiae, Canestriniidae) et Lorryia formosa n. sp. (Stomatostigmata, Tydeidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 34: 1-10. [In French]
21. Darbemamieh, M., Kaźmierski, A. and Paktinat-Saeij, S. 2020. New records and remarks on Tydeoidea (Acari: Trombidiformes) from Mazandaran province of Iran. Persian Journal of Acarology, 9 (3): 243-253.
22. de Jong, Y. 2016. Fauna Europaea. Fauna Europaea Consortium. Checklist dataset. doi: 10.15468/ymk1bx
23. Dilbaryan, K.P., Stepanyan, I.Е.E. and Stepanyan, N.T. 2022. Tydeids (Acariformes, Tydeidae) of the fauna in Armenia. In: Theory and practice of combating parasitic diseases. Indyukhova, E.N. (Ed.). Moscow, Russia, 178-183. [In Russian] doi: 10.31016/978-5-6046256-9-9.2022.23.178-183
24. Donczyk, J. 2006. Free-living Tydeoidea (Acari: Actinedida) from the vicinity of Gniew (northern Poland): preliminary report. Biological Letters, 43: 163-168.
25. Englert, W.D. 1989. Biological control of red spider Panonychus ulmi (Koch) in viticulture of the Federal Republic of Germany. In: Plant-protection problems and prospects of integrated control in viticulture. Cavalloro, R. (Ed.). Commission of the European Communities, Luxembourg, 181-186.
26. Escobar-Garcia, H.A., Andrade, D.J. de, Ueckermann, E.A. and André, H.M. 2023. A new species of the genus Brachytydeus Thor sensu André (Acari: Tydeidae) and a key to all known species from Peru. Zootaxa, 5319 (2): 263-274. doi: 10.11646/zootaxa.5319.2.7
27. Estrada-Venegas, E.G., Acuña-Soto, J.A., Chaires-Grijalva, M.P. and Equihua-Martínez, A. 2021. Mites (Arachnida: Acari). In: Diversidad Biológica. Vol. 1. Solorio, A.B. and Salgado, I.T. (Eds). Universidad Autónoma del Estado de Morelo, México, 271 pp. [In Spanish]
28. Felix, J.R. 2013. Reported incidences of parasitic infections in marine mammals from 1892 to 1978. Zea E-Books. 20, Lincon, Nebraska, USA, 149 pp.
29. Garcia-Mari, F., Marzal, C. and Laborda, R. 1985a. Tydeid species (Acari: Actinedida) and its population dynamics in Spanish citrus orchards. Boletim da Sociedade Portuguesa de Entomologia, 4 (Suppl. 1): 199-207. (Actas do II congresso Ibérico Entomologia) [In Spanish]
30. Garcia Mari, F., Laborda, R., Costa-Comelles, J., Ferragut, F. and Marzal, C. 1985b. Acaros fitofagos y depredadores en nuestros citricos. Cuadernos de Fitopatologia, 2: 54-63. [In Spanish]
31. Garcia Mari, F., Rivero, J.M. del, Marzal, C., Costa-Comelles, J., Laborda, R. and Ferragut, F. 1985c. Acaros fitofagos y depredadores en nuestros citricos. Cuadernos de Fitopatologia, 2: 132-137. [In Spanish]
32. Garman, P. 1948. Mite species from apple trees in Connecticut. Connecticut Agricultural Experiment Station Bulletin, 520: 1-27.
33. Gerson, U. 1968. Five tydeid mites from Israel (Acarina: Prostigmata). Israel Journal of Zoology, 17: 191-198.
34. Ghasemi, A. and Hajizadeh, J. 2021. Fauna and identification key for prostigmatid mites (Acari: Prostigmata) associated with greenhouses in Rasht city. Plant Pest Research, 11 (1): 1-14. [In Persian]
35. Gupta, S.K. 1992. Arachnida: Plant mites (Acari). In: State fauna Series. 3. Fauna of West Bengal, Part 3: 61-211.
36. Gupta, S.K. 2002. A monograph on plant inhabiting predatory mites (Part 1). Orders: Prostigmata, Astigmata and Cryptostigmata. Memoirs of the Zoological Survey of India, 19 (2): 1-183.
37. Hajiqanbar, M.R., Hadad Irani Nejad, K. and Talebi Chaichi, P. 2002. Mites associated with sugar beet foliage in Miandoab (East Azerbaijan). Proceedings of the 15th Iranian Plant Protection Congress, 7-11 September 2002, Kermanshah, Iran, 234-235.
38. Iberfauna. 2005. Species Lorryia mali. (Oudemans, 1929). In: IBERFAUNA. El Banco de Datos de la Fauna Ibérica. Museo Nacional de Ciencias Naturales (CSIC). Available from: https://iberfauna.mncn.csic.es/showficha.aspx ?rank=T&idtax=79550 (Last accessed: 20 July 2025). [In Spanish]
39. Jalilirad, M., Hajizadeh, J. and Noei, J. 2013. Fauna of prostigmatic mites (Acari: Prostigmata) associated with citrus orchards in Guilan Province. Journal of Plant Pests Research, 2 (4): 1-13.
40. Kamali, K., Ostovan, H. and Atamehr, A. 2001. A catalog of the mites and ticks (Acari) of Iran. Islamic Azad Scientific Publication centre, Iran, 196 pp.
41. Kammerer, C.F. 2006. Notes on some preoccupied names in Arthropoda. Acta Zootaxonomica Sinica, 31: 269-271.
42. Kaźmierski, A. 1981 (1980). Description of Venilia longina sp. nov. (Acari, Actinedida: Tydeidae) with a key to all species of the genus. Bulletin de l'Académie Polonaise des Sciences, 28 (10-11): 647-652.
43. Kaźmierski, A. 1989. Revision of the genera Tydeus Koch sensu André, Homeotydeus André and Orthotydeus André with description of a new genus and four new species of Tydeinae (Acari: Actinedida: Tydeidae). Mitteilungen Hamburgisches Zoologisches Museum und Institut, 86: 289-314.
44. Kaźmierski, A. 1990. Tydeidae mites (Actinedida, Acari) of the Świętokrzyskie Mountains. Fragmenta Faunistica, 33: 181-189. [In Polish] doi: 10.3161/00159301FF1990.33.12.181
45. Kaźmierski, A. 1996. A revision of the subfamilies Pretydeinae and Tydeinae (Acari, Actinedida: Tydeidae). Part II. The subfamily Pretydeinae André, 1979  New taxa, species, review, key and considerations. Mitteilungen Hamburgisches Zoologisches Museum und Institut, 93: 171-198.
46. Kaźmierski, A. 1998. Tydeinae of the world: generic relationships, new and redescribed taxa and keys to all species. A revision of the subfamilies Pretydeinae and Tydeinae (Acari: Actinedida: Tydeidae)  Part IV. Acta Zoologica Cracoviensia, 41: 283-455.
47. Kaźmierski, A. 2008a. Three new tydein species of the genus Brachytydeus Thor, 1931 sensu André 2005 (Acari: Actinedida: Tydeidae) from Siberia. Annales Zoologici, 58 (2): 347-355. doi: 10.3161/000345408X326663
48. Kaźmierski, A. 2008b. Description of two new species of Tydeinae (Acari: Actinedida: Tydeidae) from Spain with the remarks about the Iberian species of subfamily. Annales Zoologici, 58: 357-363. doi: 10.3161/000345408X326672
49. Kaźmierski, A. 2009. Three new Tydeinae species (Acari: Actinedida: Tydeidae) from Poland. Annales Zoologici, 59: 107-117. doi: 10.3161/000345409X432619
50. Kaźmierski, A., Marciniak, M. and Sikora, B. 2018. Tydeinae mites (Acariformes: Prostigmata: Tydeidae) from bird nests with description of three new species. Systematic and Applied Acarology, 23 (5): 803-823. doi: 10.11158/saa.23.5.3
51. Kaźmierski, A. and Panou, H.N. 1997. Lorryia reticuloinsignia, a new species of tydeid mite from the Balkan Peninsula (Acari: Prostigmata). Genus, 8: 727-734.
52. Khaustov, A.A. 2014. Obituary: Nikolay Nikolayevich Kuznetsov. Acarina, 22 (2): 161-166.
53. Khaustov, A.A. 2023. Contribution to the fauna of Tydeidae (Acari: Prostigmata) from Western Siberia, Russia. Acarologia, 63 (2): 491-521. doi: 10.24349/7fyj-me9u
54. Khaustov, A.A. and Khaustov, V.A. 2023. The first faunistic data on Tydeidae (Acari: Prostigmata) from the Altay, Russia. Acarina, 31 (2): 249-270. doi: 10.21684/0132-8077-2023-31-2-249-270
55. Kulczycki, A.G. 1992. Peculiarities of plant dwelling tydeid mite distribution (Acariformes, Tydeidae) in Kanev Nature Reserve and its Buffer Zone. Vestnik Zoologii, 5: 50-56. [In Russian]
56. Kulczycki, A.G. 1993. Habitat and plant biomorph preference in three cosmopolite Tydeus mite species (Acariformes, Tydeidae) of the forest-steppe Black Sea Nature Reserve. Vestnik Zoologii, 1993 (4): 48-51. [In Russian]
57. Kulikova, L. 2007. Influence of different types of gardens on plant-borne mite communities in Moldova. Revistă Stiintifică Universitatii de Stat din Moldova, 2 (7): 45-49. [In Russian]
58. Kulikova, L. 2008. Acarofauna in different types of habitat in the territory of the Republic of Moldova. Revistă Stiintifică Universitatii de Stat din Moldova, 2 (12): 111-118. [In Russian]
59. Kulikova, L. 2010. Diversity of distribution of plant-inhabiting ticks in forests and on adjacent grain crops. Revistă Stiintifică Universitatii de Stat din Moldova, 1 (31): 134-138. [In Russian]
60. Kulikova, L. 2011. Mites of fruit plantations of the republic of Moldova. Oltenia. Studii şi comunicări. Ştiinţele Naturii, 27 (1): 55-62.
61. Kulikova, L. 2014. Distribution of higher plant mites in the “Codru” and “Căpriana–Scoreni” reserves of the Republic of Moldova. Simpozionul" Sustainable use and protection of animal world diversity. Chișinău, Moldova, 30-31 October 2014. Institutul de Zoologie al AŞM, Chisinau, Moldova, 158-159. [In Russian]
62. Kulikova, L. 2015. Diversity of mites (Arachnida: Acariformes et Parasitiformes) on the leaves of wild trees and shrubs from the forests of the Republic of Moldova. Oltenia. Studii şi comunicări. Ştiinţele Naturii, 31 (2): 75-82.
63. Kulikova, L. 2016. Mites (Parasitiformes and Acariformes) of trees from landscape reserves of the Republic of Moldova. Journal of Wetlands Biodiversity, 6: 167-173.
64. Kulikova, L. 2021. Analysis of the mite fauna of woody and shrub plants of the Dniester river and its tributaries. Academician Leo Berg  145: Collection of Scientific Articles, Academician Leo Berg  145: Collection of Scientific Articles, Eco-TIRAS, Bendery: 140-145. [In Russian]
65. Kuznetsov, N.N. 1972. Mites of the genus Paralorryia (Tydeidae) from the Crimea. Zoologicheskiy Zhurnal, 51 (1): 28-35. [In Russian]
66. Kuznetsov, N.N. 1974. A contribution to the fauna of the family Tydeidae (Acariformes) of the Central-Chernozem District. Zoologicheskiy Zhurnal, 53 (7): 1092-1093. [In Russian]
67. Kuznetsov, N.N. 1975. New genus and species of Tydeidae (Acariformes) of the Crimean fauna. Zoologicheskiy Zhurnal, 54 (8): 1255-1257. [In Russian]
68. Kuznetsov, N.N. 1979. On revision of the family Tydeidae (Acariformes). Zoologicheskiy Zhurnal, 58 (9): 1413-1415. [In Russian]
69. Kuznetsov, N.N. and Livshitz, I.Z. 1973a. New and little known species of mites (Acariformes, Tydeidae) of the Crimea fauna. Nauchnye Doklady Vysshei Shkoly Biologicheskie Nauki, 3: 13-18. [In Russian]
70. Kuznetsov, N.N. and Livshitz, I.Z. 1973b. Some new species of mites (Acariformes, Tydeidae) of the Crimea fauna. Nauchnye Doklady Vysshei Shkoly Biologicheskie Nauki, 8: 13-18. [In Russian]
71. Kuznetsov, N.N. and Livshitz, I.Z. 1973c. New species of mites (Acariformes, Tydeidae) from the Nikitsky Botanical Gardens. Zoologicheskiy Zhurnal, 52 (2): 280-282. [In Russian]
72. Kuznetsov, N.N. and Livshitz, I.Z. 1973d. Five new species of mites from the family Tydeidae (Acariformes) from Crimea and Georgia. Zoologicheskii Zhurnal, 52 (3): 436-439. [In Russian]
73. Kuznetsov, N.N. and Livshitz, I.Z. 1973e. Three new species of the genus Paralorryia (Acariformes, Tydeidae) from the Nikitsky Botanical Gardens. Zoologicheskiy Zhurnal, 52 (4): 604-606. [In Russian]
74. Lin, J.-Z. and Zhang, Z.-Q. 2010. Tydeoidea of China: a review of progress, with a checklist. Zoosymposia, 4 (1): 51-56. doi: 10.11646/zoosymposia.4.1.4
75. Livshitz, I.Z. and Kuznetzov, N.N. 1973. Five new species of mites from the family Tydeidae (Acariformes) from Crimea and Georgia. Zoologicheskij Zhurnal, 52 (3): 436-439. [In Russian]
76. Livshitz, I.Z., Kuznetzov, N.N. and Zapletina, V.P. 1972. New species of the family Tydeidae (Acariformes) from Crimea and Azerbaijan. Zoologicheskij Zhurnal, 51: 1578-158. [In Russian]
77. Livshitz, I.Z., Kuznetzov, N.N. and Zapletina, V.P. 1973 New species of Tydeidae (Acariformes) in the fauna of Caucasus and Crimea. Zoologicheskij Zhurnal, 52: 1250-1252. [In Russian]
78. Meyer, M.K.P. and Ryke, P.A.J. 1959. New species of mites of the families Tydeidae and Labidostommidae (Acarina: Prostigmata) collected from South African plants. Acarologia, 1: 408-420.
79. Momen, F.M. and Lundqvist, L. 1995. The genus Tydeus (Acari, Prostigmata, Tydeidae) in Southern Sweden; six new species. Acarologia, 36 (1): 41-56.
80. Momen, F. and Lundqvist, L. 1996. Corticolous mites; new and unrecorded species of the genus Tydeus (Acari: Prostigmata: Tydeidae) and a key to species of Southern Sweden. Acarologia, 37 (2): 83-96.
81. Momen, F.M. and Lundqvist, L. 2005. The genera Metalorryia and Tydeus (Acari: Prostigmata: Tydeidae), new and unrecorded species from south Sweden. International Journal of Acarology, 31: 225-236. doi: 10.1080/01647950508684425
82. Monjarás-Barrera, J.I., Chacón-Hernandez, J.C., Silva, G.L. Da, Johann, L., Silva, O.S. Da, Landeros-Flores, J., Vanoye-Eligio, V., Reyeszepeda, F. and Ferla, N.J. 2019. Mites associated to chile piquín (Capsicum annuum L. var. glabriusculum) in two protect natural areas in Northeastern México. Systematic and Applied Acarology, 24 (12): 2537-2551. doi: 10.11158/saa.24.12.17
83. Mouna, M. 1998. Étude nationale sur la biodiversité. Invertébrés terrestres. Observatoire National de l’Environment du Maroc. Rabat, Morocco, 132 pp. [In French]
84. Mumford, B.C. 1965. List of intercepted plant pests, 1964. USDA, Maryland, USA.
85. Musayeva, Z.Y. and Aslanov, O.X. 2019. Predatory mites of vineyards of the Ganja-Gazakh region (Acari). Həyat elmləri və biotibb jurnalı, 74 (2): 52-57. [In Azeri]
86. Oudemans, A.C. 1928. Acarologische aanteekeningen XCIV. Entomologische Berichten 7: 374-382 [516]. [In Dutch]
87. Oudemans, A.C. 1929a. Acarologische aanteekeningen XCV. Entomologische Berichten, 7: 393-399 [518]. [In Dutch]
88. Oudemans, A.C. 1929b. Acarologische Aanteekeningen XCVIII. Entomologische Berichten, 7: 476-485 [525]. [In Dutch]
89. Oudemans, A.C. 1937. Kritisch historisch Overzicht der Acarologie. Derde Gedeelde, Band C. Leiden: E. J. Brill. p I-XXIII, 799-1348. [566]. [In Dutch] doi: 10.1163/9789004629813
90. Panou, H.N. and Emmanouel, N.G. 1995. New records of tydeid mites from Greece, with description of Lorryia brachypous sp. nov. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 11 (152): 211-220.
91. Rasmy, A.H. and El-Bagoury, M.E. 1980 (1979). A new species of genus Paralorryia from Egypt (Acarina: Tydeidae). Acarologia, 21: 194-196.
92. Ripka, G., Fain, A., Kaźmierski, A., Kreiter, S. and Magowski, W.Ł. 2002. Recent data to the knowledge of the arboreal mite fauna in Hungary (Acari: Mesostigmata, Prostigmata, and Astigmata). Acarologia, 42 (3): 271-281. doi: 10.1556/aphyt.40.2005.1-2.13
93. Ripka, G., Király, G., Szabó, Á. and Kaźmierski, A. 2022. Recent additions to the plant-inhabiting mite fauna of three European countries (Acari: Acariformes: Tydeoidea, Stigmaeidae). Systematic and Applied Acarology, 27 (5): 865-887. doi: 10.11158/saa.27.5.4
94. Ripka, G., Szabó, Á., Tempfli, B. and Varga, M. 2013. New plant-inhabiting mite records from Hungary (Acari: Mesostigmata, Prostigmata and Astigmata) II. Acta Phytopathologica et Entomologica Hungarica, 48: 237-244. doi: 10.1556/aphyt.48.2013.2.5
95. Sadeghi, H., Łaniecka, I. and Kaźmierski, A. 2012. Tydeoid mites (Acari: Triophtydeidae, Iolinidae, Tydeidae) of Razavi Khorasan Province, Iran, with description of three new species. Annales Zoologici, 62: 99-114. doi: 10.3161/000345412X633685
96. Sadeghi Namaghi, H. 1995. Survey of mites (Acari) fauna of pomaceous fruit trees in Mashhad region. Agricultural Science and Technology Journal, 9 (1): 110-120.
97. Schiess, T. 1981. Neue Tydeidenarten (Acari, Actinedida, Tydeidae) aus einem alpinen Rasen (Caricetum firmae, 2500 m) des Schweizer Nationalparkes. Entomologica Basiliensia, 6: 78-107. [In German]
98. Sepasgosarian, H. 1997. The world genera and species of the family Tydeidae (Actinedida: Acaridia). Journal of the Entomological Society of Iran, supplement 7: 1-54. [Part in Persian].
99. Silva, G.L. da, Metzelthin, M.H., Silva, O.S. da and Ferla, N.J. 2016. Catalogue of the mite family Tydeidae (Acari: Prostigmata) with the world key to the species. Zootaxa, 4135 (1): 1-68. doi: 10.11646/zootaxa.4135.1.1
100. Sirvid, P.J., Zhang, Z.-Q., Harvey, M.S., Rhode, B.E., Cook, D.R., Bartsch, I. and Staples, D.A. 2011. 6: Phylum ARTHROPODA CHELICERATA: horseshoe crabs, arachnids, sea spiders. In: New Zealand inventory of biodiversity. Vol. 2. Kingdom Animalia. Chaetognatha, Ecdysozoa, ichnofossils. Gordon, D.P. (Ed.). Canterbury University Press, Christchurch, New Zealand, 50-89.
101. Soysal, M. and Akyazı, R. 2018. Mite species of the vegetable crops in Ordu province with first report of Amblyseius rademacheri Dosse, 1958 (Mesostigmata: Phytoseiidae) in Turkey. Türkiye Entomoloji Dergisi, 42 (4): 265-286. doi: 10.16970/entoted.447218
102. Stepanyan, N.T. 2023. A contribution to the fauna of Tideidae mites (Arcariformes, Tydeidae) of Armenia. Biological Journal of Armenia, 2-3 (75): 144-149. doi: 10.54503/0366-5119-2023.75.2-3-144
103. Stepanyan, N. and Zarikian, N. 2024. Exploring Tydeoidea mites in Armenia. Euroasian Entomological Journal, 23 (4): 215-219. doi: 10.15298/euroasentj.23.04.05
104. Stojnić, B., Panou, H., Papadoulis, R., Petanović, R. and Emmanouel, N. 2002. The present knowledge and new records of phytoseiid and tydeid mites (Acari: Phytoseiidae, Tydeidae) for the fauna of Serbia and Montenegro. Acta Entomologica Serbica, 7 (1/2): 111-117.
105. Tempfli, B., Pénzes, J., Fail, J. and Szabó, Á. 2015. The occurrence of tydeoid mites (Acari: Tydeoidea) in Hungarian vineyards. Systematic and Applied Acarology, 20 (8): 937-954. doi: 10.11158/saa.20.8.9
106. Thor, S. 1931. Norwegische Tydeidae. I-VII. Mit Kennzeichnung vier neuer Gattungen. Zoologischer Anzeiger, 94: 89-10. [In German]
107. Thor, S. 1933. Tydeidae, Ereynetidae. Das Tierreich, 60: 1-84. [In German]
108. Timár, E., Bozai, J. and Bürgés, G. 2004. Additions to the knowledge of mites living on garlic. Növényvédelem, 40 (1): 17-25.
109. Tseng, Y.-H. 1985. A taxonomical study of the Tydeidae from Taiwan (Acarina, Trombidiformes) (Part 1). Journal of the Taiwan Museum, 38 (1): 129-167.
110. Ueckermann, E.A., Çobanoğlu, S. and Sağlam, H.D. 2019. Re-description of two new tydeid records (Acari: Trombidiformes) with a key to Tydoidea species of Turkey. Systematic and applied Acarology, 24 (3): 497-450. doi: 10.11158/saa.24.3.13
111. Ueckermann, E.A. and Grout, T.G. 2007. Tydeoid mites (Acari: Tydeidae, Edbakerellidae, Iolinidae) occurring on citrus in southern Africa. Journal of Natural History, 41: 2351-2378. doi: 10. 1080/00222930701589921
112. Ueckermann, E.A. and Smith Meyer, M.K.P. 1979. African Tydeidae (Acari). 2. The genus Paralorryia Baker, 1965. Phytophylactica, 11: 117-127.
113. Ueckermann, E.A. and Smith Meyer, M.K.P. 1988. South African Acari. IV. Some mites of the Addo Elephant National Park. Koedoe, 31: 31-51. doi: 10.4102/koedoe.v31i1.483
114. Vacante, V. 1985. Olive tree mites in Sicily  A first paper. In: Integrated pest control in olive-groves. Cavalloro, R. and Crovetti, A. (Eds). Published for the Commission of the European Communities by Balkema, A.A., Rotterdam, The Netherlands, 285-299.
115. Willmann, C. 1952. Die Milbenfauna der Nordseeinsel Wangerooge. Veröffenlichungen des Institute für Meeresforschung in Bremerhaven, 1: 139-186, plts 26-28. [In German]
116. Womersley, H. 1937: Acarina. Scientific reports of the Australasian Antarctic Expedition, 1911-1914. Series C - Zoology and Botany, 10 (6): 1-24.
117. Zacarias, M.S. and Moraes, G.J. de. 2002. Mite diversity (Arthropoda: Acari) on euphorbiaceous plants in three localities in the state of São Paulo. Biota Neotropica, 2: 1-12. doi: 10.1590/S1676-06032002000200004
118. Zaher, M.A. and Elbagoury, M.E. 1981. New tydeid mite, Paralorryia bakeri n. sp. from Egypt (Prostigmata: Tydeidae). Acarologia, 12 (2): 179-180.